Research

Research

Research on salamanders: systematics, ecology, and management.
The southeastern U. S. is the worldwide center of familial diversity and probable geographical region where salamanders evolved. Florida, having 27 species, is abundantly endowed with salamanders and yet they still are poorly studied. Little life history and other ecological information is available about them, but they are very sensitive ecological indicators and should be thoroughly studied and monitored. An indication of the poor state of knowledge about this important group of animals is the fact that there are at least five additional species of salamanders in Florida that have not been formally recognized and are new to science. One of CPI's most important research projects is to rectify the problem of lack of knowledge about these very important animals. Following is some of the progress we have made and are underway with in this regard:

• Amphiuma pholeter. Several research papers have been published on this species' biology and a first-draft manuscript has been completed on its life history, ecology, and food habits.

• Notophthalmus perstriatus. A ten-year drift fence study on the life history, ecology, and distribution of this rare and declining species was funded by the U. S. Fish and Wildlife Service, U. S. Forest Service, and Florida Department of Transporation. A large monograph on this project is in preparation, to include five years of data on larval phenology of the Striped Newt in temporary ponds, and hydrological analysis of some 26 ponds in which the Striped Newt and Gopher Frog breed. Altogether, some 30 species of amphibians and reptiles utilize ponds in the Munson Sandhills where the study has been underway. An alarming decline of the use of Munson Sandhills ponds by the Striped Newt has been documented. Possible causes of the decline and suggestions for management of the sandhills will be discussed.

• Ambystoma cingulatum. CPI has published several important papers on the biology of this species. Based upon our work and that of others, the flatwood salamander in 1999 was listed by the US Fish and Wildlife Service as a threatened species. We plan a drift fence study of the immigration, emigration, and larval ecology and phenology in selected temporary ponds on the Apalachicola National Forest. In addition, we plan to study the use of longleaf pine terrestrial habitat by the metamorphs and adults, and the population response of the species to prescribed burning at different seasons.

• Desmognathus auriculatus. Vital research on this species' biology and conservation has been completed and several important research papers and manuscripts have been generated, including the SSAR Catalogue Account and "Contributions on the ecology of the Southern Dusky Salamander, Desmognathus auriuclatus (Holbrook) in Florida, Georgia, and Alabama." In 1998 field work supported by a grant from Eglin Air Force Base lead to the discovery that the southern dusky salamander appears to have declined dramatically since the mid-1970s and has been extirpated from many parts of its original geographic range (Virginia to east Texas). This is the first amphibian in the eastern U.S. for which a range-wide, unexplained decline has been documented and the first salamander to merit declining amphibian status similar to that of many frogs. An important research paper about this is scheduled to be published in the March issue of Southeastern Naturalist, entitled "Declines in ravine-inhabiting dusky salamanders of the southeastern U. S. Coastal Plain." Research is continuing on this problem.

• Desmognathus. Although CPI has published many research papers on this genus in Florida and adjacent states, there still is some work to do to thoroughly understand the systematic relationships of this genus in Florida. Having described the new species, Desmognathus apalachicolae, in 1989, apparently there is at least one other undescribed species still in the Florida panhandle. Biochemical and morphological research and field work are underway to clarify the status of this genus.

• Eurycea quadridigitata complex. Field work by CPI biologists in the past two years has revealed that there are three species masquerading under the same name in Florida. FSU graduate student Kenny Wray is working with Bruce Means and other members of his Ph. D. committee to elucidate the phylogenetic relationships of this large complex of populations from east Texas to North Carolina.  Kenny is using the techniques of DNA analysis of these populations to acquire evolutionary insights.

• A comprehensive technical monograph on the "Salamanders of Florida" or "Salamanders of the southeastern U. S. Coastal Plain" is sorely needed. Over the years since 1968, B. Means has amassed a tremendous amount of information on the biology of Coastal Plain salamanders, including an alcohol preserved collection of about 20,000 specimens. This is a major project that CPI would like to seek funding for and complete in the next five years.

• Effects of fire on amphibians and reptiles.

  • In 1982 Means and Campbell published the only review entitled, "Effects of prescribed burning on amphibians and reptiles." This paper was buried in an obscure symposium (Prescribed fire and wildlife in Southern forests of the Belle W. Baruch Forest Science Institute, Clemson University, Clemson, South Carolina) and should be updated with 16 years of new data and published in a more visible outlet. A worldwide literature search through 1994 has been completed, and further review and a publication is planned for the near future.

• Vertebrate Food Web Relationships of Southeastern US Large Snakes.
  A major research goal of the Coastal Plains Institute is to study the ecology of all the large snakes of the longleaf pine ecosystem and determine their importance in the larger food web of the all the vertebrates. The nine species of snakes are the gray rat snake (Elaphe obsoleta), red rat snake (E. guttata), coachwhip (Masticophis flagellum), black racer (Coluber constrictor), pine snake (Pituophis melanoleucus), indigo snake (Drymarchon corais), eastern kingsnake (Lampropeltis getulus), eastern diamondback rattlesnake (Crotalus adamanteus), and cottonmouth (Agkistrodon piscivorus). Unfortunately, the indigo snake has been largely extirpated from panhandle Florida. Snakes may have a much greater role in the population dynamics of other vertebrates (such as ground nesting birds and small mammals, for instance) than wildlife biologists realize. With densities of up to several of these predators per acre in many cases, these large snakes have been unassessed in their potential positive and negative impacts. Snakes take rats that feed upon the eggs of ground-nesting birds, for instance, but on the other hand kingsnakes and ratsnakes feed upon eggs, themselves, and upon birds. Moreover, kingsnakes and indigo snakes feed upon snakes that feed upon other vertebrates. What sorts of ecological responses have taken place following the extirpation of the indigo snake, how do snakes affect ground-nesting birds, and dozens more questions are the focus of our studies. Some of our progress follows:

  • Eastern Diamondback Rattlesnake (Crotalus adamanteus)

    • A long-term study of the world's largest rattlesnake was undertaken in 1976 on Tall Timbers Research Station in northern Leon County, Florida. Research on this species has been supported by the Coastal Plains Institute since 1984. Presently a flatwoods population is under radiotelemetry study near our Sumatra property in Liberty County, Florida, and it is planned to inaugurate a study on a population inhabiting a mature beech/magnolia forest.

    • The write-up of a 20-year study of the life history and ecology is close to being completed as a book-length monograph to be entitled "Diamonds in the rough: natural history of the eastern diamondback rattlesnake." CPI is presently seeking a publisher.

    • Habitat utilization, home range, population density, and the effects of collecting are being assessed on Little St. Simons Island, Georgia. Funding is through CPI and some is being sought through the Georgia DNR.

    • Habitat utilization, home range, and population density in flatwoods habitat is underway utilizing radiotelemetry in Compartment 100 on the Apalachicola National Forest.

  • Cottonmouth (Agkistrodon piscivorus) From 1976-1984, a population of the cottonmouth was studied in the Red Hills on Tall Timbers Research Station utilizing radiotelemetry. CPI now has Coastal Lowlands populations in flatwoods and river swamps under study near its facility in the Apalachicola National Forest, also using radiotelemetry. Results of all these studies are planned for a book on the species' ecology.

  • Eastern kingsnake (Lampropeltis getulus) o Culminating research on the populations of kingsnake across the Florida Panhandle, a research paper entitled, "Pattern variation in the kingsnake, Lampropeltis getulus, across the Apalachicola Region of Florida," has been published and is entitled, Biogeography and pattern variation of kingsnakes, Lampropeltis getula, in the Apalachicola region of Florida.   You can access this research paper by clicking on this URL: http://www.calacademy.org/research/herpetology/ch/ch/2001/5/index.htm.  Drs. Kenneth Krysko and Walter Judd published a follow-up second paper that can be read here (PDF file).

  • Ratsnakes and the remaining large species o Limited knowledge about the other snakes has been accumulating. Unfortunately, the research that was underway on Tall Timbers Research Station was cut short in 1984. However, plans to resume it in the near future are underway pending funding.

• Grant funds are being sought to do a controlled study of the use by vertebrates of root cavities formed by rotting and/or burned out bases of longleaf pines and other trees.

1. Vertebrate Paleoecology of the Aucilla River Basin.
   In the late 1960s through the early 1980s, B. Means worked the bottom sediments of the Aucilla and Wacissa rivers for vertebrate fossils. Following that time, he and Harley Means and Ryan Means have accumulated a large data set of the occurrence of living vertebrates in the same drainage basin, which heads up in the Red Hills region of southern Georgia and flows through the Florida Big Bend. Presently CPI is working up all this material in the attempt to do a paleoecological study comparing the present-day vertebrate fauna with that of the recent past (through the end of the Pleistocene about 10,000 years ago). This study will compare all the vertebrates, small and large, that have lived in this drainage basin over the past 15,000 years and try to advance some reasonable hypotheses about 1) what impacts the extinctions have had on the present flora and fauna, and 2) how and why have the surviving species been able to survive.

2. Wiregrass restoration:Results of a 17-year transplant experiment. In 1980 B. Means transplanted wiregrass, Aristida aristida, onto burn-study plots on Tall Timbers Research Station. In the summer and fall of 1999 Trina Mitchell surveyed the site and mapped all the new plants that have recruited there in the past 20 years. Tom Ostertag is working with Bruce Means to do a second follow-up census and publish the results. This will be the first research paper to describe the results of wiregrass establishment over such a long time period. Research paper planned for submission in 2000.

3. Small Isolated Water Bodies (Temporary Ponds).
   Twenty-five to thirty-five percent of the amphibians and reptiles living in any longleaf pine forest are obligately dependent upon small isolated water bodies (temporary ponds) in their life cycles. Beginning in 1968, B. Means has seined and dipnetted temporary ponds widely in the north Florida-south Georgia area for the purpose of determining the life cycles of many of the vertebrates using such water bodies. Then, in 1994, Means initiated long-term intensive studies of 250 temporary ponds in the Munson Sandhills physiographic region south of Tallahassee Florida. This research is funded by 1) the U. S. Forest Service, 2) the U. S. Fish and Wildlife Service, and 3) the Florida Department of Transportation. Three subprojects of the larger study are:

   • To work out the breeding cycles of the gopher frog (Rana capito) and striped newt (Notophthalmus perstriatus) in the Munson Sandhills. Both species of amphibians are under consideration by the U. S. Fish and Wildlife Service for threatened status.

   • To determine the seasonal and annual use of Study Pond #1 by 31 species of amphibians and reptiles, and to assess the impacts on their populations of U. S. Highway 319 adjacent to the eastern side of the pond.

   • To determine whether certain silvicultural practices are harmful to the biodiversity of the Munson Sandhills.

4. Land Conservancy and Regional Biodiversity Conservation Activities

   • Sumatra property longleaf pine/seepage bog restoration
     In 1994 the Coastal Plains Institute acquired a superb 80-acre in-holding in the Apalachicola National Forest about 3 miles NW of Sumatra, Liberty County, Florida. Unfortunately, the dry uplands (about 40 acres) had been bedded and planted to slash pine about 1980, but our tract is surrounded by the very best stand of second-growth longleaf pine/wiregrass (by the Forest Service's own reckoning, Compartments 98 & 100) left on the national forest. In addition, it has 3 small (2-5 acres) seepage bogs, joins a 150-acre wet flat, and has a blackwater stream running through it. It is our goal to fully restore this site to its natural conditions, and to build a small facility there to serve as our biological research station and environmental education center.

   • Perdido Bay/Crown Pointe Preserve
     In 1996, the Coastal Plains Institute acquired a superb 170-acre white-topped pitcher plant bog on the eastern shores of Perdido Bay in Escambia County, Florida (west of Pensacola). We acquired this property as a mitigation package that allowed a developer to create a subdivision on about 20 acres of wetlands in exchange for giving CPI the ownership and a small endowment to assist in our management over time. CPI signed a conservation easement with the Florida Department of Environmental Protection in which our mission is to restore and maintain in perpetuity the wet flat to its presettlement condition. Mainly, this means applying prescribed fire frequently to suppress the evergreen shrubs (especially titi) that have invaded it. We have already divided the property into 40-acre management compartments and prescribe burned them for the first time in 1997. Coastal Plains Institute is looking for more such tracts to own and manage. We are available to receive mitigation lands, and looking for prospects.

SCIENTIFIC AND TECHNICAL PUBLICATIONS, CONTRACT REPORTS, POPULAR ARTICLES, AND BOOKS 

BY D. BRUCE MEANS

• Means, D. Bruce and Clive J. Longden.  1970.  Observations on the occurrence of Desmognathus monticola in Florida.  Herpetologica 26(4):396-399. 

• Means, D. Bruce.  1971.  Dentitional morphology in desmognathine salamanders (Amphibia: Plethodontidae).  Association of Southeastern Biologists Bulletin 18(2):45.

• Means, D. Bruce.  1972.  Notes on the autumn breeding biology of Ambystoma cingulatum    (Cope) (Amphibia: Urodela: Ambystomatidae).  Association of Southeastern Biologists Bulletin 19(2):84.  

• Means, D. Bruce.  1972.  Osteology of the skull and atlas of Amphiuma pholeter Neill (Amphibia: Urodela: Amphiumidae).  Association of Southeastern Biologists Bulletin 19(2):84.  

• Hobbs, Horton H., Jr. and D. Bruce Means.  1972.  Two new troglobitic crayfishes (Decapoda, Astacidae) from Florida.  Proceedings of the Biological Society of Washington 84(46):393-410.

• Means, D. Bruce.  1972.  Comments on undivided teeth in urodeles.  Copeia 1972(3):386-388.

• Means, D. Bruce.  1974.  The status of Desmognathus brimleyorum Stejneger and an analysis of the genus Desmognathus in Florida.  Bulletin of the Florida State Museum, Biological Sciences, 18(1):1-100.

• Means, D. Bruce.  1974.  City of Tallahassee Powerline Project: Faunal Impact Study.  Report under contract with the City of Tallahassee, Florida, 198 pages.

• Means, D. Bruce.  1974.  A survey of the amphibians, reptiles and mammals inhabiting St. George Island, Franklin County, Florida with comments on vulnerable aspects of their ecology.  21 pages in R. J. Livingston and N. M. Thompson, editors.   Field and laboratory studies concerning effects of various pollutants on estuarine and coastal organisms with application to the management of the Apalachicola Bay system (North Florida, U.S.A.).  Florida Sea-grant project.

• Means, D. Bruce.  1975.  Competitive exclusion along a habitat gradient between two species of salamanders (Desmognathus) in western Florida.  Journal of Biogeography 2(4):253-263.

• Means, D. Bruce.  1975.  Environmental sketch of Leon County, Florida with special emphasis on vertebrates.  Report under contract with Leon County, Florida, 119 pages.  

• Means, D. Bruce.  1976.  Survey of the status of amphibians and reptiles of the Apalachicola National Forest, Florida.  Report under contract with the U. S. Forest Service, 60 pages.

• Means, D. Bruce.  1976.  Survey of the status of amphibians and reptiles of the Osceola National Forest, Florida.  Report under contract with the U. S. Forest Service, 52 pages.  

• Means, D. Bruce.  1976.  Survey of the status of amphibians and reptiles of the Ocala National Forest, Florida.  Report under contract with the U. S. Forest Service, 43 pages.

• Means, D. Bruce.  1976.  Evolutionary ecology studies on salamanders of the genus Desmognathus.  Part I: Competitive exclusion along a habitat gradient between two species of salamanders (Desmognathus) in western Florida; Part II: Life history, growth and body size variation in populations of a streamside salamander (Desmognathus brimleyorum) on adjacent mountains.  Ph. D. thesis, Florida State University, Tallahassee, Florida.  [Dissertations Abstracts International 36(8):76-2676.]

• Means, D. Bruce and Clive J. Longden.  1976.  Aspects of the biology and zoogeography of the pine barrens treefrog (Hyla andersonii) in northern Florida.  Herpetologica 32(2):117-130.

• Means, D. Bruce.  1976.  Geographic distribution: Heterodon nasicus gloydi.  Herpetological Review (3):123.

• Means, D. Bruce.  1976.  Endangered species: Pine barrens treefrog.  Florida Naturalist 49(5):15-21.  Text plus seven photographs including cover in color.

• Means, D. Bruce.  1976.  Review of Mount, R. H.  The Reptiles and Amphibians of Alabama.  Association of Southeastern Biologists Bulletin 23(1):13.   

• Abele, Lawrence G. and D. Bruce Means.  1977.  Sesarma jarvisi and S. cookei: Montane, terrestrial grapsid crabs in Jamaica.  Crustaceana 32(1):91-93.

• Means, D. Bruce.  1977.  Aspects of the significance to terrestrial vertebrates of the Apalachicola River drainage basin, Florida.  Florida Marine Research Publications 26:23-67.

• Means, D. Bruce.  1978.  Endangered pine barrens treefrog (Hyla andersonii Baird), pages 3-4; Rare one-toed amphiuma (Amphiuma pholeter Neill), pages 6-7; Rare seal salamander (Desmognathus monticola Dunn), pages 7-9; Rare Georgia blind salamander (Haideotriton wallacei Carr), pages 9-11; Rare four-toed salamander (Hemidactylium scutatum (Schlegel)), pages 11-12; Rare mole snake (Lampropeltis calligaster rhombomaculata (Holbrook)), pages 58-60; Rare Apalachicola populations of the eastern common kingsnake including L. g. goini (Lampropeltis getulus (Linnaeus)), pages 60-61; Status undetermined coal skink (Eumeces anthracinus pluvialis Cope), pages 73-74.  In Roy McDiarmid, editor.  Rare and endangered biota of Florida, Volume 3, Amphibians and reptiles.  University of Florida Press, Gainesville, Florida.  USA. 

• Means, D. Bruce and S. P. Christman.  1978.  Rare carpenter frog, Rana virgatipes Cope.  Pages 15-17 in R. McDiarmid, editor.  Rare and endangered biota of Florida, Volume 3, Amphibians and reptiles.  University of Florida Press, Gainesville, Florida, USA.

• Christman, S. P. and D. Bruce Means.  1978.  Rare striped newt, Notophthalmus perstriatus (Bishop).  Pages 14-15 in R. McDiarmid, editor.  Rare and endangered biota of Florida, Volume 3, Amphibians and reptiles.  University of Florida Press, Gainesville, Florida, USA.

• Means, D. Bruce.  1978.  Investigations into the decline of the Wakulla River Limpkin population 1975-1978.  Report under contract with the Edward Ball Wildlife Foundation, 11 pages.

• Means, D. Bruce, and P. E. Moler.  1979.  The pine barrens treefrog: fire, seepage bogs, and management implications.  Pages 77-83 in R. R. Odum and L. Landers, editors.  Proceedings of the Rare and Endangered Wildlife Symposium, August 3-4, Athens, Georgia.  Georgia Department of Natural Resources, Game and Fish Division Technical Bulletin WL-4.  

• Means, D. Bruce.  1980.  Quail research at Tall Timbers: rising to the challenge.  Pages 28-31 in S. A. Vezey and Roy Komarek, editors.  Proceedings of the Tall Timbers Quail Management Seminar.  Held 5 March 1980 at Thomasville, GA. 

• Means, D. Bruce.  1980.  Stewardship plan for Phipps Point Nature Preserve.  Report under contract with The Nature Conservancy, 75 pages, (with R. L. Crawford).  

• Means, D. Bruce.  1981.  Steepheads: Florida's little-known canyon lands.  Enfo, December 1981:1-4.  Text plus two b/w photographs.

• Karlin, A. A., D. Bruce Means, S. I. Guttman and D. Lambright.  1982.  Systematics and the status of Hyla andersonii (Anura:Hylidae) in Florida.  Copeia 1982(3):176-178.

• Means, D. Bruce and Howard W. Campbell.  1982.  Effects of prescribed burning on amphibians and reptiles.  Pages 89-97 in G.  W. Wood, editor. Prescribed fire and wildlife in Southern forests.  Proceedings of a symposium.  Belle W. Baruch Forest Science Institute, Clemson University, Clemson, South Carolina, USA.

• Means, D. Bruce.  1982.  Feasibility of advanced wastewater treatment facility, Fort Polk, Louisiana.  Report under contract with the U. S. Army, 14 pages.  

• Means, D. Bruce.  1982.  Responses to winter burrow flooding of the gopher tortoise (Gopherus polyphemus Daudin).  Herpetologica 38(4):521-525.

• Means, D. Bruce and E. V. Komarek.  1983.  Forum: Ecological animal control by habitat management.  Environmental Management 7(1):1-2.

• Means, D. Bruce.  1983.  The enigmatic pine barrens treefrog.   Florida Wildlife 37(1):16-19.  Text plus eight color photographs.

• Means, D. Bruce.  1983.  Welcoming remarks.  Pages 1-4 in R. Franz and R. J. Bryant, editors.  The gopher tortoise and its sandhill habitat.  Proceedings of the Third Annual Meeting, Gopher Tortoise Council.  

• Means, D. Bruce, J. B. Atkinson, and D. G. Cook.  1983.  Overwintering behavior of two syntopic crotaline snakes (Agkistrodon piscivorus, Crotalus adamanteus) in northern Florida.  63rd annual meeting of the American Society of Ichthyologists and Herpetologists, Tallahassee, Florida.

• Means, D. Bruce.  1983.  Tall Timbers Research Station.  Pages 74-79 in Chapin, C. M., editor.   The Florida-Georgia Field Trial Association.  Princeton University Press, Princeton, New Jersey, USA.

• Brown, Lauren E. and D. Bruce Means.  1984.  Fossorial behavior and ecology of the chorus frog Pseudacris ornata.  Amphibia-Reptilia 5:261-273.

• Means, D. Bruce.  1984.  Survey of beech-magnolia forests of north Florida.  Report under contract with The Nature Conservancy, 66 pages. 

• Means, D. Bruce.  1985.  Ecological Critique of the Proposed Management Plan for Florida National Forests.  Report under contract with the National Wildlife Federation, 31 pages.  

• Means, D. Bruce.  1985.  Radio-tracking the Eastern Diamondback Rattlesnake.  National Geographic Society Research Reports 18:529-536.

• Means, D. Bruce.  1985.  Ice flowers: Discovery.  Bitter Sweet South 1(2):7-10.  Text plus seven color photographs.

• Means, D. Bruce.  1985.  Nature blooms in winter.  Bitter Sweet South 1(3):12-14.  Text plus five color photographs.

• Means, D. Bruce.  1985.  The cotton rat: Base of the food chain.  Enfo, Spring 1985:6-7.  

• Means, D. Bruce.  1985.  The canyonlands of Florida.  Nature Conservancy News, September/ October:13-17.  Text plus two color photographs.

• Means, D. Bruce and Gerald O. Grow.  1985.  The endangered longleaf pine community.  ENFO, September:1-12.  Text plus nine b/w photographs.

• Means, D. Bruce.  1985.  Delicate balance.  Florida Wildlife.  November-December, p. 37.

• Means, D. Bruce.  1986.  Threatened pine barrens treefrog, Hyla andersonii Baird, pages 29-30; Threatened dusky gopher frog, Rana areolata sevosa Goin and Netting, pages 30-31; Species of special concern flatwoods salamander, Ambystoma cingulatum Cope, pages 42-43; Species of special concern eastern diamondback rattlesnake, Crotalus adamanteus Beauvois, pages 48-49; Poorly known one-toed amphiuma, Amphiuma pholeter Neill, pages 56-57; Poorly known southern dusky salamander, Desmognathus auriculatus (Holbrook), pages 58-59; Poorly known Gulf Coast mud salamander, Pseudotriton montanus flavissimus Hallowell, page 62; Poorly known Southern red salamander, Pseudotriton ruber vioscai Bishop, pages 62-63 in Robert H. Mount, editor.  Vertebrate animals of Alabama in need of special attention.  Proceedings of the First Alabama Non-Game Conference, 15-16 July 1983, Auburn University, Alabama, i-xiv, 1-124.

• Means, D. Bruce.  1986.  Keynote address: John Muir's walk through the Southeast--one hundred seventeen years later.  Pp. 1-3 in D. R. Jackson and R. J. Bryant (eds.).  The gopher tortoise and its community.  Proc. 5th Ann. Mtg., Gopher Tortoise Council, Gainesville, Florida. pp. i-vii, 1-93.

• Means, D. Bruce.  1986.  Alligator, Blacksnake, Coral Snake, Crocodile, Garter Snake, Rattlesnake, Water Moccasin, King Snake, Terrapin: Accounts in World Book   Encyclopedia, 7th ed.

• Means, D. Bruce.  1986.  Rattlesnake fever: it’s contagious!  The Skimmer 2(2):8.  Text plus one photograph. 

• Means, D. Bruce.  1986.  Life History and Ecology of the Eastern Diamondback Rattlesnake.  Report under contract with Florida Game and Fresh Water Fish Commission, project number GFC-84-13, 331 pages. 

• Means, D. Bruce.  1986.  Ecological Reasons for an Appeal of Forest Service Proposed Management Plan for Florida National Forests.  Report under contract with the National Wildlife Federation, 23 pages.  

• Means, D. Bruce.  1986.  The Status of the Indigo Snake.  Report under contract with Tidewater II Properties, Okaloosa-Walton counties, Florida, 9 pages. 

• Means, D. Bruce.  1986.  Developing a Methodology for Surveying the Ordinary High Water Line: Is It Feasible?  Report under contract with the Florida Department of Natural Resources, 128 pages. 

• Means, D. Bruce and Daniel I. Simberloff.  1987.  The peninsula effect: habitat-correlated species decline in Florida's herpetofauna.  Journal of Biogeography 14(4):551-568.

• Means, D. Bruce.  1987.  Impact of current forestry practices on national forests in Florida.  Report under contract with The Wilderness Society, 59 pages.  

• Means, D. Bruce.  1987.  A brief history of OHWL determinations in Florida and listing of pertinent literature.  In David Gibson (ed.).  Development of Procedural Methodology to Enable Florida Land Surveyors to Determine the Ordinary High Water Line Boundary of Sovereign Lands.  Final report under contract to Florida Institute of Government, Star Project 86-043, 340 pages.  

• Means, D. Bruce.  1987.  The application of hydrology to the determination of the ordinary high water line.  In David Gibson (ed.), Development of procedural methodology to enable Florida land surveyors to determine the ordinary high water line boundary of sovereign lands.  Final report under contract to Florida Institute of Government, Star Project 86-043, 340 pages.  

• Means, D. Bruce.  1988.  An ecological characterization of the Florida Panhandle.  Report under contract with Florida Department of Environmental Regulation and U. S. Fish and Wildlife Service.  U.S. Fish Wild. Serv. Biol. Rept. 88(12), 277 pages.  

• Means, D. Bruce.  1988.  Ordinary High Water Primer (A Review of the Scientific and Legal Basis for OHWL Determination).  Volume A.  First Ed.  Report under Contract No. C5171 with Florida Department of Natural Resources, 192 pages. 

• Means, D. Bruce.  1988.  Ordinary High Water Workbook.  (A Manual to Guide the Surveyor in Establishing the Ordinary High Water Line on Non-tidal Sovereign Lands).  Volume B.  First Ed.  Report under Contract No. C5171 with Florida Department of Natural Resources, 94 pages. 

• Wolfe, S. H., J. A. Reidenauer, and D. B. Means.  1988.  An ecological characterization of the Florida panhandle.  U. S. Fish & Wildife Service Biological Report 88(12):1-277.

• Means, D. Bruce.  1988.  Management recommendations for the gopher tortoise in longleaf pine ecosystems.  Pp. 41-56 in C. K. Dodd, Jr. (ed.).  Gopher tortoise habitat management: strategies and options.  Proc. 6th Ann. Mtg. Gopher Tortoise Council, Florida Museum of Natural History, Gainesville.  58 pp.

• Means, D. Bruce.  1988.  Published comments as a member of summation panel for symposium entitled, “Wiregrass biology and management: Maintaining groundcover integrity in longleaf pine ecosystems.”  Co-sponsored by Coastal Plains Institute, Gopher Tortoise Council, Natural Areas Association, and KBN Engineering, Valdosta State College, October 13, 1988.

• Means, D. Bruce and Alvan A. Karlin.  1989.  A new species of Desmognathus (Amphibia: Caudata: Plethodontidae) from the eastern Gulf Coastal Plain.  Herpetologica 45(1):37-46.

• Means, D. Bruce and David S. Martin.  1989.  Mating phenologies in the genus Crotalus: evidence and implications.  Invited paper for The Biology of Pitvipers Symposium, University of Texas at Arlington, 17-19 November 1989.  

• Means, D. Bruce.  1989.  Comments of the Coastal Plains Institute submitted to the Subcommittee on Forests, Family Farms, and Energy of the Committee on Agriculture, United States House of Representatives, 15 November 1989.  Pages 195-225 in National Forest Management; Serial #101-42:195-225.  Washington, D. C.

• Means, D. Bruce.  1989.  The Casiquiare mystery.  South American Explorer #23:14-19.  Text plus four photographs.

• Means, D. Bruce.  1990.  The Case of the Two-quail rattlesnake.  Florida Wildlife 44(2):19-20.  Text plus one color photograph.

• Means, D. Bruce.  1990.  Where Angel trod: Ascent of Auyantepui.  South American Explorer #25:17-23.  Text plus two photographs.

• Means, D. Bruce.  1990.  Snakes alive.  South American Explorer #26:36-39.  Text plus two photographs including cover .

• Means, D. Bruce.  1990.  Florida wetlands.  Florida Wildlife 44(5):32-33.  Text plus one color photograph. 

• Means, D. Bruce.  1990.  Seepage bogs.  Florida Wildlife 44(5):34-37.  Text plus seven color photographs.

• Means, D. Bruce.  1990.  Temporary ponds.  Florida Wildlife 44(6):12-16.  Text plus five color photographs.

• Means, D. Bruce.  1990.  Geographic distribution: Anolis sagrei.  SSAR Herpetological Review 21(4):96.

• Means, D. Bruce.  1990.  Geographic distribution: Hemidactylus turcicus.  SSAR Herpetological Review 21(4):96.

• Means, D. Bruce.  1990.  Published comments as a member of the discussion panel on Silvicultural effects on groundcover plant communities in longleaf pine forests.   The Longleaf Pine Ecosystem: Ecology, Restoration and Management.  Tall Timbers Fire Ecology Conference, Tallahassee, Florida.  

• Means, D. Bruce.  1991.  River bottomlands.  Florida Wildlife 45(1):11-16.  Text plus six color photographs.

• Means, D. Bruce.  1991.  Florida’s steepheads: Unique canyonlands.  Florida Wildlife 45(3):25-28.  Text plus four color photographs.

• Means, D. Bruce.  1991.  The Aucilla hunt company.  North Florida Journal 1(1):2-16.

• Means, D. Bruce.  1991.  Do snakes swallow their young?  North Florida Journal 1(1):18-19.

• Means, D. Bruce.  1992.  Rio Napo misadventure, Part I.  South American Explorer 31:12-20.  Text plus six photographs.

• Means, D. Bruce.  1992.  Rattlesnake roundups in the southeastern United States: What do we know about roundup species?  Presentation made at the 1992 Annual Gopher Tortoise Council Meetings, Stephen Foster State Park, Fargo, Georgia, 23-25 October.

• Means, D. Bruce.  1992.  Rare Pine Barrens Tree Frog, pages 20-25; Rare One-toed Amphiuma, pages 34-38; Rare Seal Salamander, pages 44-48; Rare Georgia Blind Salamander, pages 49-53; Rare Four-toed Salamander, pages 54-57; Rare Southern Coal Skink, pages 219-222; Rare Mole Snake, pages 227-231; Rare Eastern Common Kingsnake, pages 232-236; Rare Southern Copperhead, pages 242-246.  In Paul E. Moler, ed.  Rare and endangered biota of Florida.  Volume III.  Amphibians and Reptiles.  University Presses of Florida, Gainesville.

• Means, D. Bruce and Steven P. Christman.  1992.  Rare Carpenter Frog.  Pages 26-29 in Paul E. Moler, ed.  Rare and endangered biota of Florida.  Volume III.  Amphibians and Reptiles. University Presses of Florida, Gainesville.

• Steven P. Christman and D. Bruce Means.  1992.  Rare Striped Newt.  Pages 62-65 in Paul E. Moler, ed.  Rare and endangered biota of Florida.  Volume III.  Amphibians and Reptiles.  University Presses of Florida, Gainesville.

• Means, D. Bruce.  1992.  Potential Sites for Native and High Quality Successional Forest in the City of Tallahassee and Leon County, Florida.  Report under subcontract on City of Tallahassee contract to Tall Timbers Research Station & Dr. Sharon Hermann, 29 pages.

• Means, D. Bruce.  1992.  Evaluation of the Effects of Large-scale Disturbance on Gulf Coastal Wetlands: Assessment of the Impacts of Hurricanes and Fires on Vegetation Dynamics of the St. Marks National Wildlife Refuge.  Report under subcontract on U. S. Fish and Wildlife Service grant to Dr. William Platt, 25 pages.  

• Means, D. Bruce.  1992.  Contributions toward an Ecological Characterization of the Mountain Pine Ridge, Belize, Central America.  Report under contract with Mt. Pleasant, Ltd., Hidden Valley Inn.  203 pages. 

• Karlin, Alvan A., Sheldon I. Guttman, and D. Bruce Means.  1993.  Population structure in the Ouachita Mountain Dusky Salamander, Desmognathus brimleyorum (Caudata: Plethodontidae).  Southwestern Naturalist 38(1):36-42.

• Means, D. Bruce.  1993.  Desmognathus apalachicolae.  Catalogue of the American Amphibians and Reptiles 556.1-556.2.

• Means, D. Bruce.  1993.  Wetlands fire ecology in the Guayana Highlands.  Presentation made to 19th Tall Timbers Fire Ecology Conference, Tallahassee, Florida, 4-5 November.

• Means, D. Bruce.  1993.  Rio Napo misadventure, Part II.  South American Explorer 32:15-21.  Text plus five photographs.

• Means, D. Bruce.  1993.  Sez who? Of waterfalls and boundary markers.  South American Explorer 34:16-22. Text plus seven photographs, including cover in color.

• Means, D. Bruce.  1993.  Diamonds in the rough, Little St. Simons Island.  The League of Florida Herpetological Societies Newsletter, October:17-24.

• Means, D. Bruce.  1993.  National Forests in Florida, a fact sheet/white paper.  Report under contract with The Nature Conservancy, 58 pages.  1993.

• Means, D. Bruce.  1993.  Environmental survey of Tyndall Air Force Base, Bay County, Florida.  Report under contract with Florida Natural Areas Inventory, The Nature Conservancy.  153 pages.  

• Means, D. Bruce. 1994.  Handbook for guides, Billie Swamp Safari; Ecology of Big Cypress Region of South Florida.  Report under contract with the Seminole Nation, 56 pages.  

• Means, D. Bruce. T. E. Ostertag, & D. Printiss.  1994.  Florida populations of the striped newt, Notophthalmus perstriatus, west of the Suwannee River.  Contributions to life history, ecology, and distribution.  I.  Report under contract with the U. S. Fish and Wildlife Service, Jackson, MS, 57 pages.  

• Means, D. Bruce, T. E. Ostertag, & D. Printiss.  1994.  Distribution, habitat ecology, and management of the striped newt, Notophthalmus perstriatus, in the Apalachicola National Forest, Florida.  Report under contract with the U. S. Forest Service, National Forests in Florida, Tallahassee, FL, 30 pages. 

• Karlin, Alvan A. and D. Bruce Means.  1994.  Genetic variation in the aquatic salamander genus Amphiuma.  American Midland Naturalist 132(1):1-9. 

• Means, D. Bruce.  1994.  Diamonds in the rough.  A “prepublication” of eight chapters from the forthcoming book, DIAMONDS IN THE ROUGH.  North Florida Journal, Vol. 2:1-28.

• Means, D. Bruce.  1994.  Second time around’s no charm.  The League of Florida Herpetological Societies Newsletter, Febuary: 21-27. 

• Means, D. Bruce.  1994.  Venomous snakes of South America.  South American Explorer 36:23-29.  Text plus five photographs.

• Means, D. Bruce.  1994.  Florida uplands.  Florida Wildlife 48(3):9.

• Means, D. Bruce.  1994.  Florida scrub.  Florida Wildlife 48(3):10-13.  Text plus two color photographs.

• Means, D. Bruce.  1994.  Longleaf pine forests.  Florida Wildlife 48(5):2-6.  Text plus five color photographs. 

• Healey, Kevin.  1994.  Map of Venezuela.  International Travel Map Productions, Vancouver, B. C.  Two color photographs including cover by D. B. Means.

• Means, D. Bruce.  1994.  Temperate hardwood forests.  Florida Wildlife 48(6):20-23.

• Means, D. Bruce.  1994.  Venomous snakes of South America.  League of Florida Herpetological Societies Newsletter, reprinted from South American Explorer.  December 1994:15-21. 

• Means, D. Bruce, John Palis, and Mary Baggett.  1994.  Effects of slash pine silviculture on Florida panhandle populations of the flatwoods salamander (Ambystoma cingulatum).  Paper read at the August joint meetings of the Herpetologist’s League and Society for the Study of Amphibians and Reptiles, University of Georgia, Athens.   

• Martin, W. H., and D. Bruce Means.  1994.  Distribution of the eastern diamondback rattlesnake (Crotalus adamanteus).  Poster presented at the August joint meetings of the Herpetologist’s League and Society for the Study of Amphibians and Reptiles, University of Georgia, Athens. 

• Means, D. Bruce.  1994.  Coastal Plains ecosystems as influenced by settlement and urban development.  Symposium on urban runoff, December 12-13, Florida State University and City of Tallahassee, FL.

• Means, D. Bruce.  1995.  Suwannee River in Okefenokee Swamp, 1 color photograph.  Florida Wildlife 49(1):10.

• Means, D. Bruce.  1995.  Fire ecology of the Guayana Region, northeastern South America.  Pages 61-77 in Susan I. Cerulean and R. Todd Engstrom, eds.  Fire in wetlands: a management perspective.  Proceedings of the Tall Timbers Fire Ecology Conference No. 19.  Tall Timbers Research Station, Tallahassee, FL.

• Means, D. Bruce.  1995.  Diel, seasonal, and spatial activity patterns of the eastern diamondback rattlesnake in the center of its range.  38th Annual Meeting of the Society for the Study of Amphibians and Reptiles.  Appalachian State University, Boone, NC.

• Means, D. Bruce.  1995.  Management of populations of the eastern diamondback rattlesnake.  38th Annual Meeting of the Society for the Study of Amphibians and Reptiles.  Appalachian State University, Boone, NC.

• Martin, W. H. and D. Bruce Means.  1995.  Distribution of the eastern diamondback rattlesnake.  38th Annual Meeting of the Society for the Study of Amphibians and Reptiles.  Appalachian State University, Boone, NC.

• Means, D. Bruce.  1995.  Longleaf pine forest, going, going....  Page 40-41 in Eastern Old-growth Forests Symposium of the 22nd Annual Natural Areas Conference, University of Arkansas, Fayetteville, AR.  25-30 October.

• Means, D. Bruce.  1995.  The value of tree bases and stumpholes as habitat for wildlife.  Page 41 in the 22nd Annual Natural Areas Conference, University of Arkansas, Fayetteville, AR.  25-30 October.

• Means, D. Bruce.  1995.  Amphibians and reptiles of Torreya State Park (with special emphasis on the Rock Creek Tract).  Report under contract with Florida Department of Environmental Protection, 70 pages.    

• Means, D. Bruce.  1995.  Element Stewardship Abstract: Eastern Diamondback Rattlesnake.  Report under contract with The Nature Conservancy, Southeast Regional Office, Chapel Hill, N. C.  11 pages.  

• Means, D. Bruce.  1995.  Conservation Strategy for Endangered Florida torreya, Torreya taxifolia.  Report under contract with The Nature Conservancy, Southeast Regional Office, Chapel Hill, N. C.  8 pages.  

• Means, D. Bruce.  1995.  Conservation Strategy for Endangered Chapman's rhododendron, Rhododendron chapmanii.  Report under contract with The Nature Conservancy, Southeast Regional Office, Chapel Hill, N. C.  11 pages. 

• Means, D. Bruce. 1995.  Conservation Strategy for Threatened Godfrey's butterwort, Pinguicula ionantha.  Report under contract with The Nature Conservancy, Southeast Regional Office, Chapel Hill, N. C.  10 pages.

• Means, D. Bruce.  1995.  Conservation Strategy for Threatened Miccosukee gooseberry, Ribes echinellum.  Report under contract with The Nature Conservancy, Southeast Regional Office, Chapel Hill, N. C.  8 pages.  

• Means, D. Bruce.  1995.  Conservation Strategy for Threatened Florida skullcap, Scutellaria floridana.  Report under contract with The Nature Conservancy, Southeast Regional Office, Chapel Hill, N. C.  10 pages. 

• Means, D. Bruce.  1996.  Longleaf pine forest: Importance to biodiversity.  Pages 12-14 in John S. Kush, ed.  Proceedings of the Conference on Longleaf Pine Forest Management by Private Landowners, Mobile, AL.

• Means, D. Bruce, John G. Palis, and Mary Baggett.  1996.  Effects of slash pine silviculture on a Florida population of flatwoods salamander.  Conservation Biology 10(2):426-437.

• Means, D. Bruce.  1996a.  Chapter 15.  Longleaf pine forest, going, going....  Pages 210-229 in Mary Byrd Davis, ed.  Eastern old-growth forest: Prospects for rediscovery and recovery.  Island Press, Washington, D. C.  

• Means, D. Bruce.  1996b.  A preliminary consideration of highway impacts on herpetofauna inhabiting small isolated wetlands in the southeastern U.S. Coastal Plain.  Pages 1-11 in G. L. Evink, P. Garrett, D. Zeigler, and J. Berry, eds.  Trends in addressing transportation related wildlife mortality.  Proceedings of the Transportation Related Wildlife Mortality Seminar, 30 April-2 May, 1996, Orlando, FL.  Florida Department of Transportation, Tallahassee, FL.

• Means, D. Bruce.  1996c.  Geographic distribution: Anolis sagrai.  SSAR Herpetological  Review 27(3):151-152.

• Means, D. Bruce.  1996d.  Geographic distribution: Gekko gecko.  SSAR Herpetological  Review 27(3):152.

• Means, D. Bruce.  1996e.  Geographic distribution: Hemidactylus turcicus.  SSAR Herpetological Review 27(3):152.

• Means, D. Bruce.  1996f.  Amphiuma pholeter.  Catalogue of the American Amphibians and Reptiles 622.1-622.2. 

• Means, D. Bruce and D. Printiss.  1996.  Larval life cycle of the striped newt, Notophthalmus perstriatus, on the Apalachicola National Forest, Florida.  Report under contract with the U. S. Forest Service, National Forests in Florida, Tallahassee, FL, 30 pages. 

• Printiss, D. J. and D. B. Means.  1996.  Distribution of the flatwoods salamander, Ambystoma cingulatum, on the Osceola National Forest, Florida.  Report under contract with the U.S. Forest Service, National Forests in Florida, Tallahassee, FL.  1996.  

• Swanson, Helge R. and D. B. Means. 1996.  Miccosukee Road Greenway: An environmentally sensitive areas analysis.  Report under contract with Trust for Public Lands, Tallahassee, FL, 9 pages.  

• Means, D. Bruce and D. J. Printiss.  1966.  Use of a temporary pond by amphibians and reptiles in the Munson Sandhills of the Apalachicola National Forest with special emphasis on the striped newt and gopher frog, Year 1:  September 1995 - September 1996.  Final report under contract with the U. S. Forest Service, National Forests in Florida, Tallahassee, FL, 38 pages.  September 1996. 

• Means, D. Bruce and T. Lewis.  1997.  Geographic distribution: Amphiuma means.  SSAR Herpetological Review 28(1):47. 

• Means, D. Bruce.  1997.  Wiregrass restoration: Probable shading effects in a slash pine plantation.  Restoration and Management Notes 15(1):53-56

• Means, D. Bruce and R. C. Means.  1997.  Use of a temporary pond by amphibians and reptiles in the Munson Sandhills of the Apalachicola National Forest with special emphasis on the striped newt and gopher frog, Year 2:  September 1996 - September 1997.  Final report under contract with the U. S. Forest Service, National Forests in Florida, Tallahassee, FL, 23 pages.  September 1997. 

• Means, D. Bruce.  1997.  Saving species on our National Forests: Case studies from the Southern Region.  Report under contract with the Defenders of Wildlife, Washington, D. C.   40 pages.  October 1997.  

• Means, D. Bruce.  1997.  Natural history of Mountain Pine Ridge, Belize.  Bull Run Overseas, Ltd., Benque Viejo del Carmen, Belize.  94 pages.

• Means, D. Bruce.  1998a.  Geographic distribution: Ambystoma cingulatum.  SSAR Herpetological Review 29(1):47. 

• Means, D. Bruce.  1998.  One full-page color and two black and white photographs in article entitled "Snakebite!" by Carolee Boyles-Sprenkel.  Florida Game and Fish 1998(1):44-51.  

• Means, D. Bruce.  1998.  Vanishing natural heritage: Wet flats; Conservationist's Notebook.  The American Gardener 77(1):20-21.

• Means, D. Bruce.  1998.  Review of Snakes in Question: The Smithsonian Answer Book by Carl H. Ernst and George R. Zug.  Copeia 1998(1):257-258. 

• Means, D. Bruce.  1998b.  Geographic distribution: Lampropeltis getula.  SSAR Herpetological Review 29(2):113. 

• Means, D. Bruce.  1998c.  Geographic distribution: Agkistrodon contortrix contortrix.  SSAR Herpetological Review 29(3):175.

• Means, D. Bruce. and Steven P. Christman.  1998.  Geographic distribution: Farancia abacura.  SSAR Herpetological Review 29(3):177.

• Means, D. Bruce.  1998.  Review of The Venomous Reptiles of Latin America by Jonathon A. Campbell and William W. Lamar.  South American Explorer 51(Spring 1998):51-52.

• Means, D. Bruce.  1998.  Foreward.  Pages x-xiii in Crescent Rivers by Todd Bertolaet.  Florida University Presses, Gainesville.

• Means, D. Bruce and R. C. Means.  1998.  Distribution of the striped newt (Notophthalmus perstriatus) and gopher frog (Rana capito) in the Munson Sand Hills of the Florida panhandle.  Final report to the U. S. Fish and Wildlife Service, Jackson, MS, for Order No. 43910-5-0077.  April 1998.  42 pages.  

• Means, D. Bruce and R. C. Means.  1998.  Red Hills Survey for breeding pond habitat of the flatwoods salamander (Ambystoma cingulatum), gopher frog (Rana capito), and striped newt (Notophthalmus perstriatus) in the Tallahassee Red Hills of Leon, Gadsden, and Jefferson counties, Florida, and the Tifton Uplands of Thomas and Grady counties, Georgia.  Final report to the U. S. Fish and Wildlife Service, Jackson, MS, for Order No. 43910-5-0091.   23 pages.  April 1998.  (By D. B. Means and R. C. Means.)

• Means, D. Bruce.  1998.  Amphibians and reptiles of Apalachicola Bluffs and Ravines Preserve, The Nature Conservancy.  Final Report to The Nature Conservancy.  25 pages.  December 1998.  

• Means, D. Bruce.  1999.  Salamanders of Eglin Air Force Base: Towards a long-term monitoring program.  Final Report to the Natural Resources Branch at Jackson Guard, Eglin AFB, for contract No. F0865197M5602.  126 pages.  March 1999.  

• Means, D. Bruce, and Anne Harvey.  1999.  Barbour's map turtle in the diet of nesting bald eagles.  Florida Field Naturalist 27(1):14-16.  

• Means, D. Bruce.  1999.  Snake charmer.  National Wildlife 37(2): 36-41.

• Means, D. Bruce.  1999.  Venomous snakes of Florida.  Florida Wildlife 53(5):13-20, plus cover.

• Means, D. Bruce.  1999b.  Desmognathus auriculatus.  Catalogue of the American Amphibians and Reptiles:681.1-681.6.

• Means, D. Bruce.  1999c.  Desmognathus brimleyorum.  Catalogue of the American Amphibians and Reptiles:682.1-682.4.

• Means, D. Bruce.  1999d.  The Effects of Highway Mortality on Four Species of Amphibians at a Small, Temporary Pond in Northern Florida.  Pages 125-128 in G. Evink, P. Garrett, and D. Zeigler, eds.  Proceedings of the Third International Conference on Wildlife Ecology and Transportation, September 13-16, 1999.  Missoula, Montana.  

• Means, D. Bruce.  2000.  Chapter 14.  Southeastern U. S. Coastal Plain habitats of the Plethodontidae: the importance of relief, ravines, and seepage.  Pages 287-302 in R. C. Bruce, R. J. Jaeger, and L. D. Houck, eds.  The Biology of Plethodontid Salamanders.  Plenum Publ. Corp., N. Y.  

• Martin, W. H. and D. Bruce Means.  2000.  Geographic distribution and habitat relationships of the eastern diamondback rattlesnake, Crotalus adamanteus.  Herpetological Natural History 7(1):9-35.

• Means, D. Bruce.  2000.  This charming snake.  BBC Wildlife Magazine 18(2):40-46.  

• Means, D. Bruce.  2000.  Non-venomous snakes of Florida.  Florida Wildlife 54(3):13-20.

• Means, D. Bruce.  2001.  Reptile.  World Book Encyclopedia, Vol. R:250-253.

• Means, D. Bruce.  2001.  Florida’s fabulous frogs.  LEEFlet 72:1.

• Means, D. Bruce.  2001.  Amphisbaenian.  World Book Encyclopedia. 

• Means, D. Bruce.  2001.  Just batty.  International Wildlife 31(5):28-29.

• Means, D. Bruce.  2001.  Living on the Ledge.  FAUNA 2(6):1-7.

• Means, D. Bruce and K. L. Krysko.  2001.  Biogeography and pattern variation in the Kingsnake, Lampropeltis getula, across the Apalachicola Region of Florida.  Contemporary Herpetology 2001(5):1-33.  http://www.calacademy.org/research/herpetology/ch/ch/2001/5/index.htm

• Means, D. Bruce.  2001.  Reducing impacts on rare vertebrates that require small isolated water bodies along U. S. Highway 319.  Final Report to the Florida Department of Transportation.  148 pages.  September 2001. 

• Means, D. Bruce.  2002.  Foreword in Miles Barton, Nigel Bean, Stephen Dunleavey, Ian Gray, and Adam White.  Wild New World, The Wildlife of Ice Age North America.  BBC Worldwide Ltd., London.

• Means, D. Bruce and Guy H. Means.  2002.  Geographic Distribution: Pseudobranchus striatus.  Herpetol. Rev. 33(4):316.

• Means, D. Bruce.  2003.  Means, D. Bruce.  2002.  King Rattler: Eastern Diamondback.  FAUNA 3(2):10-17.

• Means, D. Bruce.  2003.  Crack of Doom.  BBC Wildlife Magazine 21(3):52-55.

• Means, D. Bruce.  2003.  Where have all the gopher frogs gone?  Science World 60(2):10-13, cover. 

• Means, D. Bruce.  2003.  Biology of the southern dusky salamander, Desmognathus auriculatus. Part I.  A report to the U. S. Fish and Wildlife Service in Fulfillment of Order No. 1448-43910-2-M728A.  29 pages.  22 January 2003.

• Means, D. Bruce.  2003.  Churning the Food Chain: Frogs are food for bats in Central America, but frogs turn predator in Australia.  Bats 21(4):5-7.

• Means, D. Bruce.  2003.  Kukenan.  FAUNA 4(3):16-21.

• Means, D. Bruce.  2003.  Hunt for the Roughie.  FAUNA 4(4):16-21.  

• Means, D. Bruce.  2003.  Notes on the reproductive biology of the Alabama Red Hills Salamander (Phaeognathus hubrichti).  Contemporary Herpetology2003(3). http://www.cnah.org/ch/ch/2003/3/index.htm

• Means, D. Bruce.  2003.  Results of the drift fence survey of Study Pond #1, Apalachicola National Forest, Leon County, Florida, July 2002 – June 2003.  Final Report to U. S. Fish and Wildlife Service in satisfaction of Order No. 1448-43910-1-M718A.  21 pages.  30 November 2003.  

• Means, D. Bruce.  2004.  Nature’s hidden works of art.  Tallahassee Magazine 26(1):81-85.

• Means, D. Bruce.  2004.  Blossoms of Ice.  Natural History Magazine 113(1):36-37.

• Means, D. Bruce.  2004.  Walk Across Amazonas.  South American Explorer 74:6-12.

• Means, D. Bruce.  2004.  Gluttons of Chappell.  FAUNA 5(1):14-22. 

• Means, D. Bruce.  2004.  The loathed Cottonmouth Moccasin: Is it aggressive?  FAUNA 5(2):14-22.  

• Means, D. Bruce.  2004.  Ghosts of the Red Hills.   Chapter 7.  Pages 89-99 in Susan Cerulean, Janisse Ray, and Laura Newton (eds.).  Between Two Rivers.  Red Hills Writers Project, Tallahassee, FL.  

• Means, D. Bruce.  2004.  What is it?  Geographica Section, National Geographic Magazine 206(5): p. xxx.

• Means, D. Bruce.  2004.  Hard to Swallow.  Half page article featuring a quarter page photograph of Green Treefrog eating Little Bent-wing Bat.  WILD, the Canadian Wildlife Federation magazine for kids, Vol. 8(6):23. 

• Bailey, Mark A. and D. Bruce Means.  2004a.  Gopher Frog, Rana capito Le Conte and Mississippi Gopher Frog, Rana sevosa Goin and Netting.  Pages 15-17 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004b.  Flatwoods salamander, Ambystoma cingulatum (Cope).  Pages 19-20 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004c.  Southern Dusky Salamander, Desmognathus auriculatus (Holbrook). Pages 22-23 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004d.  Pine Barrens Treefrog, Hyla andersonii Baird.  Pages 25-26 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004e.  One-toed Amphiuma, Amphiuma pholeter Neill. Pages 28-29 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Bailey, Mark A. and D. Bruce Means.  2004f.  Red Hills Salamander, Phaeognathus hubrichti Highton. Pages 34-36 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004g.  Coal Skink, Eumeces anthracinus (Baird). Pages 58-59 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004h.  Eastern Kingsnake, Lampropeltis getula getula Linnaeus. Pages 65-66 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004i.  Florida Pine Snake, Pituophis melanoleucus mugitus (Barbour).  Pages 69-70 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004j.  Eastern Diamondback Rattlesnake, Crotalus adamanteus Beauvois. Pages 73-74 in R. E. Mirarchi, M. A. Bailey, T. M. Haggerty, and T. L. Best, eds.  Alabama Wildlife.  Vol. 3.  Imperiled amphibians, reptiles, birds, and mammals.  University of Alabama Press, Tuscaloosa, Alabama.

• Means, D. Bruce.  2004k.  Natural history notes: Rana capito (Florida Gopher Frog) defensive behavior.  Herpetological Review 35(2):163-164.  

• Means, D. Bruce, C. K. Dodd, Jr., S. A. Johnson, and J. G. Palis.  2004l.  Amphibians and fire in longleaf pine ecosystems.  Conservation Biology 18(3):1149-1153.   

• Means, D. Bruce.  2004m.  Geographic distribution: Ambystoma tigrunum.  Herpetological Review 35(4):402-403.  

• Means, D. Bruce.  2004n.  Geographic distribution: Bothriopsis taeniata.  Herpetological Review 35(4):410.

• Means, D. Bruce.  2005.  Mysterious Ice “Flowers.” American Gardener Magazine Jan/Feb:34-37.

• Means, D. Bruce.  2005. Report to the Florida Department of Transportation for Work Completed 1 July 2004 – 28 February 2005.  Monitoring Amphibians and Reptiles under FDOT Contract No. BD543, Work Order #7.  Life Cycles, Dispersal, and Critical Habitat Utilization of Vertebrates Dependent upon Small Isolated Water Bodies in the Munson Sandhills and Woodville Karst Plain, Leon County, Florida.  17 pages.  1 March 2005. 

• Means, D. Bruce.  2005.  Leviathan of Southern Rivers.  FAUNA Magazine 5(4):18-23.  

• Means, D. Bruce.  2005.  Steepheads: Florida’s Hidden Valleys.  Tallahassee Magazine May/June:84-89, 122-123.  Eight color photographs.

• Means, D. Bruce.  2005.  Rattlesnake. World Book Encyclopedia, Vol. R.

• Means, D. Bruce.  2005.  Crocodile. World Book Encyclopedia, Vol. C.

• Means, D. Bruce.  2005.  Hyla andersonii.  Pages 445-447 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.

• Palis, John G. and D. Bruce Means.  2005.  Ambystoma cingulatum.  Pages 608-609 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.

• Means, D. Bruce.  2005.  Amphiuma pholeter.  Pages 645-646 in Michael Lanoo, ed.  Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley.   

• Means, D. Bruce.  2005.  Desmognathus apalachicolae.  Pages 698-699 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.   

• Means, D. Bruce.  2005.  Desmognathus auriculatus.  Pages 700-701 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.   

• Means, D. Bruce.  2005.  Desmognathus brimleyorum.  Pages 701-703 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.   

• Means, D. Bruce and R. Bonett.  2005.  Desmognathus conanti.  Pages 705-706 in Michael Lanoo, ed.  Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.

• Means, D. Bruce.  2005.  Desmognathus fuscus.  Pages 708-710 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.

• Means, D. Bruce.  2005.  Haideotriton wallacei.  Pages 779-780 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.

• Dodd, C. K., Jr., S. A. Johnson, D. Bruce Means.  2005.  Notophthalmus perstriatus.  Pages 887-889 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.

• Means, D. Bruce.  2005.  Chapter 21. Pine silviculture.  Pages 139-145 in Michael Lanoo, ed. Amphibian declines: The conservation status of United States species.   University of California Press, Berkeley, CA.

• Means, D. Bruce and Ryan C. Means.  2005.  Chapter 7.  Effects of sand pine silviculture on pond-breeding amphibians in the Woodville Karst Plain of north Florida. Pages 56-61 in W. Meshaka and K. Babbitt, eds.  Status and conservation of Florida amphibians and reptiles.  Krieger Press, Melbourne, Florida.  

• Means, D. Bruce.  2005.  The value of dead tree bases and stumpholes as habitat for wildlife.  Chapter 9.    Pages 74-78 in W. Meshaka and K. Babbitt, eds.  Status and conservation of Florida amphibians and reptiles.  Krieger Press, Melbourne, Florida.  

• Means, D. Bruce.  2006.  Chapter 6.  Vertebrate faunal diversity in longleaf pine savannas.  Pages 155-213 in S. Jose, E. Jokela, and D. Miller, eds.  Longleaf Pine Ecosystems: Ecology, Management, and Restoration.  Springer, New York.   xii + 438 pp.

• Grant, T., D. R. Frost, J. P. Caldwell, R. Gagliardo, C. F. B. Haddad, P. J. R. Kok, D. B. Means, B. P. Noonan, W. Schargel, and W. C. Wheeler.  2006. Phylogenetic systematics of dart-poison frogs and their relatives (Anura: Athesphatanura: Dendrobatidae). Bulletin of the American Museum of Natural History 299: 1-262. 

• Means, D. Bruce.  2006.  Lizard. World Book Encyclopedia, Vol. L.  pp. 399-401.

• Means, D. Bruce.  2006.  Chameleon. World Book Encyclopedia, Vol. C. pp. 372.

• Means, D. Bruce.  2007.  Turtle.  World Book Encyclopedia, Vol. T. pp. 520-524.

• Means, D. Bruce.  2007.  Sea Turtle.  World Book Encyclopedia, Vol. S.

• Means, D. Bruce.  2007.  Caiman.  World Book Encyclopedia, Vol. C.

• Means, D. Bruce.  2007.  Pages 61-65 in J. Ray (ed.).  Moody Forest.  Wildfire Press, Baxley, GA.

• Means, D. Bruce and Joseph Travis.  2007.  Declines in ravine-inhabiting dusky salamanders of the southeastern US Coastal Plain.  Southeastern Naturalist 6(1):83-96. 

• Means, D. Bruce and Michelle Kalamandeen.  2007.  Geographic distribution.  Oxyrhopus melanogenys.  Herpetological Review 38(1):105. 

• Means, D. Bruce.  2007.  Life cycles, dispersal, and critical habitat utilization of vertebrates dependent upon small isolated water bodies in the Munson Sandhills and Woodville Karst Plain, Leon County, Florida.  Final Report to the Florida Department of Transportation for OMNI Project 010562.  79 pp.

• Means, D. Bruce and Stephen Richter.  2007.  Genetic verification of possible gigantism in a southern toad, Bufo terrestris.  Herpetological Review 38(3):297-298.  

• Means, D. Bruce and Jay M. Savage.  2007.  Three new malodorous rainfrogs of the genus Pristimantis (Anura: Brachycephalidae) from the Wokomung Massif in west-central Guyana, South America.  Zootaxa (1658):39-55.  

• Means, D. Bruce.  2007.  Geographic distribution.  Atractus zidoki.  Herpetological Review 38(4):484-485. 

• Stapleton, S. P., K. J. Sash, D. B. Means, W. E. Palmer, and J. P. Carroll.  2008.  Eastern kingsnake (Lampropeltis g. getula) population decline in North Florida and South Georgia.  Herpetological Review 39(1):33-35.

• Means, D. Bruce.  2008.  One-toed Amphiuma (Amphiuma pholeter).  Pages 151-153 in J. B. Jensen, C. Camnp, W. Gibbons, and M. Elliott, eds.  Amphibians and Reptiles of Georgia.  University of Georgia Press, Athens, Georgia.  575 pp.

• Means, D. Bruce.  2008.  Apalachicola Dusky Salamander (Desmognathus apalachicolae). Pages 163-165 in J. B. Jensen, C. Camp, W. Gibbons, and M. Elliott, eds.  Amphibians and Reptiles of Georgia.  University of Georgia Press, Athens, Georgia.  575 pp.

• Means, D. Bruce.  2008.  Southern Dusky Salamander (Desmognathus auriculatus). Pages 166-168 in J. B. Jensen, C. Camnp, W. Gibbons, and M. Elliott, eds.   Amphibians and Reptiles of Georgia.  University of Georgia Press, Athens, Georgia.  575 pp.

• Means, D. Bruce.  2008.  Dwarf Salamanders (Eurycea quadridigitata complex). Pages 198-201 in J. B. Jensen, C. Camp, W. Gibbons, and M. Elliott, eds.    Amphibians and Reptiles of Georgia.  University of Georgia Press, Athens, Georgia.  575 pp.

• Means, D. Bruce.  2008.  Georgia Blind Salamander (Haideotriton wallacei).  Pages 208-210 in J. B. Jensen, C. Camnp, W. Gibbons, and M. Elliott, eds.    Amphibians and Reptiles of Georgia.  University of Georgia Press, Athens, Georgia.  575 pp.

• Krysko, Kenneth L. and D. Bruce Means.  2008.  Mole Kingsnake (Lampropeltis calligaster rhombomaculata). Pages 359-360 in J. B. Jensen, C. Camnp, W. Gibbons, and M. Elliott, eds.    Amphibians and Reptiles of Georgia.  University of Georgia Press, Athens, Georgia.  575 pp.

• Means, D. Bruce.  2008.  Eastern Diamondback Rattlesnake (Crotalus adamanteus). Pages 430-432 in J. B. Jensen, C. Camnp, W. Gibbons, and M. Elliott, eds.   Amphibians and Reptiles of Georgia.  University of Georgia Press, Athens, Georgia.  575 pp.

• Means, D. Bruce, Ryan. C. Means, and Rebecca Pharr Meegan Means.  2008.  Geographic Distribution.  Anolis sagrei.  Herpetological Review 39(3):366.  

• Means, D. Bruce, William E. Duellman, and Valerie C. Clark.  2008.  Ovipositing behavior in the egg-brooding frog Stefania ayangannae (Hemiphractidae). Phyllomedusa 7(2):143-148.  

• Heinicke, Matthew P., William E. Duellman, Linda Trueb, D. Bruce Means, Ross D. MacCulloch, and S. Blair Hedges.  2009.  A new frog family (Anura: Terrarana) from South America and an expanded direct-developing clade revealed by molecular phylogeny.  Zootaxa 2211:1-35.

• Means, D. Bruce.  2009.  Effects of rattlesnake roundups on the eastern diamondback rattlesnake (Crotalus adamanteus).  Herpetological Conservation Biology 4(2):132-141.

• Magalhães, Célio, Michael Türkay, and D. Bruce Means.  2009. The status of Kunziana Pretzmann, 1971 (Crustacea: Decapoda: Pseudothelphusidae), with a redescription of the holotype of K. irengis Pretzmann, 1971.  Zootaxa 2276:40-48.

• Means, D. Bruce.  2009.  Darwin's Eureka Moment. Florida State University Research in Review. Summer 2009:40-43.

• Means, D. Bruce.  2009.  The biology of the southern dusky salamander, Desmognathus auriculatus. Part II.  Report to the U. S. Fish and Wildlife Service in Fulfillment of Order No. 1448-43910-3-M008A.  38 pages.  6 June 2009. 

• Means, D. Bruce.  2010.  Ophidism in the Green Palmsnake, Philodryas viridissimus.  Wilderness and Environmental Medicine 21:46-49.

• Means, D. Bruce.  2010.  Blocked-flight aggressive behavior in snakes.  IRCF Reptiles and Amphibians 17(2):76-78.

• Means, D. Bruce, Kenneth R. Sims, and Kenneth L. Krysko. 2010. Geographic Distribution.  Drymarchon couperi.  Herpetological Review 41(3):380. 

• Means, D. Bruce.  2010.  Time to end rattlesnake roundups: the ecological impacts of a long tradition.  The Wildlife Professional, Winter 2010:64-67.

• Kok, Philippe J. R., D. Bruce Means, and Franky Bossuyt.  2011.  A new highland species of Pristimantis Jiménez de la Espada, 1871 (Anura: Strabomantidae) from the Pantepui Region, northern South America.  Zootaxa 2934:1-19.

• Kok, Philippe J. R., Ross D. MacCulloch, D. Bruce Means, K. Roelants, I. Van Bocxlaer, and Franky Bossuyt.  2012.  Low genetic diversity in tepui summit vertebrates.  Current Biology 22:589-590, with supplementary material at doi:10.1016/j.cub.2012.06.034. 

• Kok, Philippe J. R., Ross D. MacCulloch, D. Bruce Means, K. Roelants, I. Van Bocxlaer, and Franky Bossuyt.  2012. Unlocking the secrets of a Lost World.  Science 337:1020.

• Brown, Donald J., John T. Baccus, D. Bruce Means, and Michael R. J. Forstner.  2011. Potential positive effects of fire on juvenile amphibians in a Southern USA pine forest. Journal of Fish and Wildlife Management 2(2):135-145. 

• Means, D. Bruce.  2011.  Biodiversity Dependent Upon Wetland Ecosystems Created by Seepage from the Surficial Aquifer in Bay and Washington Counties, Florida.  Final Report to The Northern Trust Company as the Sole Trustee of the James L. Knight Charitable Term Trust.  23 August 2011.  51 pages.

• Means, R. C., R. P. M. Means, D. L. Miller, M. J. Gray, S. A. Johnson, D. B. Means, and R. Brenes.  2011.  A Conservation Strategy for the Imperiled Striped Newt (Notophthalmus perstriatus) in the Apalachicola National Forest, Florida.  First Annual Report to the U. S. Forest Service, Tallahassee, Florida, September 2011.  20 pages. 

• Means, D. Bruce.  2012.  Tijd om de ratelslangenround-ups te stoppen: De ecologische gevolgen van een lange traditie.  Litteratura Serpentium 32(4):187-194.

• Means, D. Bruce.  2012.  De diamantratelslang, Zachtaardige reus onder de ratelslangen. Litteratura Serpentium 32(4):195-200.

• Kok, Philippe J. R., Bert Willaert, and D. Bruce Means.  2013.  A new diagnosis and description of Anomaloglossus roraima (La Marca, 1998) (Anura: Aromobatidae: Anomaloglossinae), with description of its tadpole and call.  South American Journal of Herpetology 8(1):1-17.

• Jungfer, K., J. Faivovich, D. B. Means, et al.  2013.  Systematics of an Amazonian puzzle: A phylogenetic analysis of spiny-backed treefrogs (Osteocephalus) reveals its paraphyly and a high number of cryptic species (Amphibia, Anura, Hylidae).  Zoologica Scripta 1-30, 1-68 (Supplementary information.)

• Kok, Philippe J. R. and D. Bruce Means.  2013.  First record of the genus Anadia (Reptilia, Squamata, Gymnophthalmidae) from Guyana based on an enigmatic specimen from Mount Kopinang, Wokomung Massif.  Boletim do Museu Paraense Emilio Goeldi. Ciencias Naturais 8(1):27-39.

• Valentine, J. and D. Bruce Means.  2014.  Florida Magnificent Wilderness, State Lands, Parks, and Natural Areas.  Portfolio, The Magazine of Lifestyles, Interest, and Leisure.  January 2014:42-46.

• Noss, Reed, William J. Platt, Bruce Sorrie, Alan Weakley, D. Bruce Means, Jennifer Costanza, and Robert Peet. 2014. How global biodiversity hotspots may go unrecognized: Lessons from the North American Coastal Plain. Diversity and Distributions 1-9, John Wiley & Sons, Ltd.

• Means, D. Bruce.  2015.  Petition to list the Southern Dusky Salamander (Desmognathus auriculatus) as a threatened species under the Endangered Species Act of 1973.  Submitted to the U. S. Secretary of the Interior.  40 pp.

• Kok, Philippe J. R., Sebastian Ratz, Marco Tegelaar, D. Bruce Means, Fabien Aubret, and Franky Bossuyt.  2015.  Out of taxonomic limbo: A name for the species of Tepuihyla (Anura: Hylidae) from the Chimantá Massif, Pantepui region, northern South America.  Salamandra, German Journal of Herpetology 51(4):283-314.

• Means, D. Bruce. 2016. Desmognathus apalachicolae (Apalachicola Dusky Salamander) and Agkistrodon contortrix contortrix (Southern Copperhead). Prey/Predator. Herpetological Review 47(1):100.

• Kok, Philippe J. R, Valerio G. Russo, Sebastian Ratz, D. Bruce Means, Ross D. MacCulloch, Amy Lathrop, Fabien Aubret, and Franky Bossuyt.  2017. Evolution in the South American ‘Lost World’: Insights from multilocus phylogeography of stefanias (Anura, Hemiphractidae, Stefania). Journal of Biogeography 44:170-181.

• Wray, K. P., D. Bruce Means, and S. J. Steppan.  2017.  Revision of the Eurycea quadridigitata (Holbrook 1842) complex of dwarf salamanders (Caudata: Plethodontdidae: Hemidactylinae) with a description of two new species.  Herpetological Monographs 31:18-46.

• Tanaka, Tara and D. Bruce Means.  2017.  Notophthalmus viridescens (Eastern newt). Predation. Herpetological Review 48(1):155-157.

• Means, D. Bruce, Jennifer Lamb, and Joseph Bernardo.  2017.  A new species of dusky salamander (Amphibia: Plethodontidae: Desmognathus) from the eastern Gulf Coastal Plain of the United States and a redescription of D. auriculatus.  Zootaxa 4263(3):467-506.

• Kok, Philippe J. R., S. Ratz, R. D. MacCulloch, A. Lathrop, R. Desfoulian, F. Aubret, and D. Bruce Means.  2017.  Historical biogeography of the paleoendemic toad genus Oreophrynella (Amphibia: Bufonidae) sheds a new light on the origin of the Pantepui endemic terrestrial biota.  Journal of Biogeography 2017:1-11.  https://doi.org/10.1111/jbi.13093

• Kok, Philippe J. R., R. Dezfoulian, D. Bruce Means, C. L. Barrio-Amoros, and A. Fouquet.  2018.  Amended diagnosis and redescription of Pristimantis marmoratus (Boulenger, 1900) (Amphibia: Craugastoridae), with description of its advertisement call and notes on its breeding ecology and phylogenetic relationships. European Journal of Taxonomy 397:1-30. 

• Kok, Philippe J. R., M. A. Bittenbinder, J. K. van der Berg, S. Marques-Souza, P. M. Sales Nunes, A. Laking, M. Teixeira Jr., A. Fouquet, D. B. Means, R. D. MacColloch, and M. Trefaut Rodrigues.  2018.  Integrative taxonomy of the gymnophthalmid lizard Neusticurus rudis Boulenger, 1900, identifies a new species in the eastern Pantepui region, north-eastern South America.  Journal of Natural History, pp. 1-38.

• Means, D. Bruce.  2018.  If Mr. Crouch Could See Me Now.  Florida Humanities Council Forum Magazine, Spring Issue:10-11. 

• Pinheiro, P. D. P., P. J. R. Kok, B. P. Noonan, D. B. Means, C. F. B. Haddad, J. Faivovich.  2018.  A new genus of Cophomantini, with comments on the taxonomic status of Boana liliae (Anura, Hylidae). Zoological Journal of the Linnaean Society, 2018, XX, 1-20.  With 6 figures.

• Means, D. Bruce and M. G. Aresco. 2019. Amphiuma pholeter.  Pages 86-87 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Desmognathus apalachicolae.  Pages 88-90 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Desmognathus auriculatus.  Pages 90-92 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Desmognathus cf. conanti.  Pages 92-94 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Desmognathus monticola.  Pages 94-97 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Eurycea hillisi.  Pages 100-101 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Eurycea quadridigitata.  Pages 101-103 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Eurycea sphagnicola.  Pages 104-105 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce. 2019. Crotalus adamanteus.  Pages 569-573 in K. L. Krysko, K. M. Enge, and P. E. Moler.  The Amphibians and Reptiles of Florida.  University of Florida Press, Gainesville, FL, USA.  

• Means, D. Bruce and César Barrio-Amorós.  2019.  Chapter 13.  Islands in the Sky, Snakes on South American Tepuis.  Pages 310-329 in H. B. Lillywhite and M. Martins, eds.  Islands and Snakes.  Oxford University Press, New York.  

• Means, D. Bruce and Chace R. Holzheuser.  2019.  Geographic Distribution.  Regina septemvittata.   Herpetological Review 50(4):749.

• Kok, P. J. R., M. A. J. Van der Velden, D. B. Means, S. Ratz, I. Josipovic, M. Boone, and R. W. McDiarmid.  2020.  Coping with the extremes: Comparative osteology of the tepui-associated toad Oreophrynella and its bearing on the evolution of osteological novelties in the genus. Zoological Journal of the Linnaean Society 2020, XX, 1-26, 11 figures.

• Holzheuser, Chace R., D. Bruce Means, and Richard Franz.  2020. Geographic Distribution. Eurycea cirrigera.  Herpetological Review 51(1):68.

• Means, D. Bruce, Chace R. Holzheuser, and David A. Beamer.  2020.  Geographic Distribution.  Eurycea sphagnicola.   Herpetological Review 51(1):68. 

• Means, D. Bruce and Margaret Gunzburger Aresco.  2020.  Natural History of the One-toed Amphiuma, Amphiuma pholeter. Herpetological Conservation and Biology 15(3):666-674.

• Warwick, A. R., L. N. Barrow, M.L. Smith, D. B. Means, A. R. Lemmon, and E. M. Lemmon.  2021.  Signatures of northeastern expansion and multiple refugia: Genomic phylogeography of the Pine Barrens Treefrog (Hyla andersonii).  Biological Journal of the Linnean Society 2021, XX, 1–15 + 10 pp supplemental.

• Laucevicious, Anthony M., Jr., Kevin M. Robertson, D. Bruce Means, and Trina R. Mitchell.  2021.  Reproduction, expansion, and growth of Aristida stricta (Wiregrass) tussocks in a transplanted population: Results of a 37-year study.  Restoration Ecology 05 April 2021. https://doi.org/10.1111/rec.13404

• Holzheuser, Chace R. and D. Bruce Means.  2021.  Decline of the Seal Salamander, Desmognathus monticola, from Florida, USA.  Herpetological Conservation and Biology 16(3):506-512.

• Holzheuser, Chace, Means, Bruce, Lips, Karen, Talley, Brooke. 15 July 2021.  Filling the data gaps to assess the population status and management of the Southern Dusky Salamander (Desmognathus auriculatus) in Florida. Final Report under FWC Agreement No. 16002 to the Florida Fish and Wildlife Conservation Commission, Tallahassee, Florida. Pp. 37 + Appendices A-G.

• Means, D. Bruce, Matthew Heinicke, S. Blair Hedges, Ross MacCulloch, and Amy Lathrop.  2023.  Exceptional diversity of  Pristimantis landfrogs  (Anura: Terraranae) on the Wokomung Massif, Guyana, with descriptions of three new species. Journal of Vertebrate Biology (Accepted for publication 3 May 2023).

• Kok, Philippe J.R. and D. Bruce Means.  2023.  Hidden in the Mists, Molecular phylogenetic position and description of a new genus and species of dipsadid snake (Serpentes, Dipsadidae, Xenodontinae) recently collected in the remote cloud forest of the Lost World. Zoological Society of the Linnean Society (submitted 01-12-2023).

• Holzheuser, Chace R., Anna Longa, D. Bruce Means, Brooke Talley, and Karen R. Lips.  2023.  Environmental DNA Detects Populations of the Southern Dusky Salamander (Desmognathus auriculatus), a Declining Amphibian in Florida.  [In preparation.  To be submitted to Herpetological Conservation Biology.]

• Means, D. Bruce, Ross MacCulloch, Amy Lathrop, and Philippe P. J. Kok.  2023.  Amphibians and reptiles of the Wokomung Massif, Guyana.  In preparation for submission to Zootaxa.

• Means, D. Bruce and Philippe P. J. Kok.  2023. Amphibians and reptiles of the Upper Paikwa River drainage basin, Guyana.  In preparation.

BOOKS

Whitney, Ellie, D. Bruce Means, and Ann R. Rudloe.  2004.  PRICELESS FLORIDA: Natural Ecosystems and Native Species.  i-viii + 423 pp.  Pineapple Press, Sarasota, FL. A book about the ecology of Florida from the Gulf of Mexico’s abyssal plain to the highest, driest, longleaf pine/turkey oak sandhills.   

Valentine, James and D. Bruce Means.  2006.  FLORIDA, Magnificent Wilderness.  Pineapple Press, Sarasota, FL.  145 pp.  A large-format photography book about Florida’s state parks and public lands.  

Means, D. Bruce.  2008.  STALKING THE PLUMED SERPENT and Other Adventures in Herpetology.  A book featuring Means’s firsthand research and adventure experiences with snakes, frogs, salamanders, earthworms and other things that go bump in the night.  Foreward by E. O. Wilson.  Pineapple Press, Sarasota, FL.  i-xi, 238 pp.

Whitney, Ellie and D. Bruce Means.  2014.  FLORIDA’S UPLANDS, Volume 1 in Florida’s Natural Ecosystems and Native Species Series.  Pineapple Press, Sarasota, FL.  166 pp. 

Whitney, Ellie, D. Bruce Means, and Anne Rudloe.  2014.  FLORIDA’S WETLANDS, Volume 2 in Florida’s Natural Ecosystems and Native Species Series.  Pineapple Press, Sarasota, FL  142 pp.  

Whitney, Ellie and D. Bruce Means, and Anne Rudloe.  2014.  FLORIDA’S WATERS, Volume 3 in Florida’s Natural Ecosystems and Native Species Series.  Pineapple Press, Sarasota, FL  166 pp. 

Valentine, James and D. Bruce Means.  2015.  FLORIDA’S MAGNIFICENT WATER.  Volume 1 in Magnificent Florida Series.  Pineapple Press, Sarasota, FL.  64 pp. 

Valentine, James and D. Bruce Means.  2015.  FLORIDA’S MAGNIFICENT LAND.  Volume 2 in Magnificent Florida Series.  Pineapple Press, Sarasota, FL.  64 pp.

Valentine, James and D. Bruce Means.  2015.  FLORIDA’S MAGNIFICENT COAST.  Volume 3 in Magnificent Florida Series.  Pineapple Press, Sarasota, FL.  64 pp.

Means, D. Bruce.  2017.  DIAMONDS IN THE ROUGH, Natural history of the Eastern Diamondback Rattlesnake.  Tall Timbers Research Station Miscellaneous Publication No. 20, Tallahassee, FL. A book on the natural history of the eastern diamondback rattlesnake based on the author’s eight-year radiotelemetry study of a free-ranging population.  Tall Timbers Press, Tallahassee, FL.  i-xxvi, 1-390.

Means, D. Bruce.  2023.  BASECAMP IN THE TROPICS, My Adventures Discovering Biodiversity in South America.  ECO Publishing, Rodeo, New Mexico.  A book about seven expeditions Means made to one of the world’s last unexplored wildernesses.  It is about natural treasures of frogs and other wildlife new to science discovered by Means on National Geographic Society funded treks.  (In press with publication date of June 2023.)

Means, D. Bruce and James Valentine.  2023.  ISLANDS IN THE SKY: Lost Worlds of El Dorado.  A book about the natural history of one of the planet's premier wilderness areas as revealed through the author's explorations and photography of remote mesas in the rainforests of the upper Orinoco basin.  (Searching for a publisher.)

Means, D. Bruce.  2023.  A NATURALIST IN THE GARDEN OF EDEN.  A book about Means’s 50-year research odyssey in the Florida panhandle where the greatest number of U. S. species occurs in frogs, snakes, turtles, carnivorous plants, ecosystems, and other groups of organisms.  The focus is on the amazing biogeography of this small, but most-biodiverse, region in the US and Canada as seen through Means’s lifelong researches and experiences.  (In preparation.)